Hi Alex,
Thanks so much for the file, as this is something that shows all the settings. Yes, if I take the file as shared, I have the same problem.
If you check your frame range in the Cinema 4D project and compare those with the Ae range, you will see that something is off. It seems the Ae numbers are the same times two.
Which lead me to check the FPS entries (plural!). There is one in the Project Setting and one in the Render Settings: Yours showed 30 [project] and 60 [render settings].
So, I adjusted the Attribute Manager> Mode> Project Settings to 60 and saved it newly (with a 60-60 name extension to be sure).
This file then showed the same frame numbers in Cinema 4D as well as in Ae. Now the keyframes in Ae line up as you would like to have them (I have to guess here, as I do not have the footage, but that will be a quick check for you).
All the best